QTL mapping and candidate gene analysis of ferrous iron and zinc toxicity tolerance at seedling stage in rice by genome-wide association study by Jian Zhang, Kai Chen, Yunlong Pang, Shahzad Amir Naveed, Xiuqin Zhao, Xiaoqian Wang, Yun Wang, Michael Dingkuhn, Julie Pasuquin, Zhikang Li, Jianlong Xu

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