Treacle controls the nucleolar response to rDNA breaks via TOPBP1 recruitment and ATR activation by Mooser, C, Symeonidou, I, Leimbacher, P, Ribeiro, A, Shorrocks, A, Jungmichel, S, Larsen, S, Knechtle, K, Jasrotia, A, Zurbriggen, D, Jeanrenaud, A, Leikauf, C, Fink, D, Nielsen, M, Blackford, A, Stucki, M