Spatial patterns of large African cats: a large-scale study on density, home range size, and home range overlap of lions Panthera leo and leopards Panthera pardus by Nams, V, Parker, DM, Weise, FJ, Bauer, H, Loveridge, A, Trethowan, P, du Preez, BD, Macdonald, D, Oriol-Cotteril, A