Collective force generated by multiple biofilaments can exceed the sum of forces due to individual ones

Collective dynamics and force generation by cytoskeletal filaments are crucial in many cellular processes. Investigating growth dynamics of a bundle of N independent cytoskeletal filaments pushing against a wall, we show that chemical switching (ATP/GTP hydrolysis) leads to a collective phenomenon t...

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Main Authors: Dipjyoti Das, Dibyendu Das, Ranjith Padinhateeri
Format: Article
Language:English
Published: IOP Publishing 2014-01-01
Series:New Journal of Physics
Subjects:
Online Access:https://doi.org/10.1088/1367-2630/16/6/063032
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author Dipjyoti Das
Dibyendu Das
Ranjith Padinhateeri
author_facet Dipjyoti Das
Dibyendu Das
Ranjith Padinhateeri
author_sort Dipjyoti Das
collection DOAJ
description Collective dynamics and force generation by cytoskeletal filaments are crucial in many cellular processes. Investigating growth dynamics of a bundle of N independent cytoskeletal filaments pushing against a wall, we show that chemical switching (ATP/GTP hydrolysis) leads to a collective phenomenon that is currently unknown. Obtaining force-velocity relations for different models that capture chemical switching, we show, analytically and numerically, that the collective stall force of N filaments is greater than N times the stall force of a single filament. Employing an exactly solvable toy model, we analytically prove the above result for N = 2. We, further, numerically show the existence of this collective phenomenon, for $N\geqslant 2$ , in realistic models (with random and sequential hydrolysis) that simulate actin and microtubule bundle growth. We make quantitative predictions for the excess forces, and argue that this collective effect is related to the non-equilibrium nature of chemical switching.
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spelling doaj.art-5336aa262e4e433d890e503fdc6508e22023-08-08T11:29:35ZengIOP PublishingNew Journal of Physics1367-26302014-01-0116606303210.1088/1367-2630/16/6/063032Collective force generated by multiple biofilaments can exceed the sum of forces due to individual onesDipjyoti Das0Dibyendu Das1Ranjith Padinhateeri2Department of Physics, Indian Institute of Technology, Bombay , Powai, Mumbai-400 076, IndiaDepartment of Physics, Indian Institute of Technology, Bombay , Powai, Mumbai-400 076, IndiaDepartment of Biosciences and Bioengineering, Indian Institute of Technology , Bombay, Powai, Mumbai-400 076, IndiaCollective dynamics and force generation by cytoskeletal filaments are crucial in many cellular processes. Investigating growth dynamics of a bundle of N independent cytoskeletal filaments pushing against a wall, we show that chemical switching (ATP/GTP hydrolysis) leads to a collective phenomenon that is currently unknown. Obtaining force-velocity relations for different models that capture chemical switching, we show, analytically and numerically, that the collective stall force of N filaments is greater than N times the stall force of a single filament. Employing an exactly solvable toy model, we analytically prove the above result for N = 2. We, further, numerically show the existence of this collective phenomenon, for $N\geqslant 2$ , in realistic models (with random and sequential hydrolysis) that simulate actin and microtubule bundle growth. We make quantitative predictions for the excess forces, and argue that this collective effect is related to the non-equilibrium nature of chemical switching.https://doi.org/10.1088/1367-2630/16/6/063032non-equilibrium chemical switchingdynamics of biofilamentscollective force generationactin and microtubules87.16.A-87.16.Ka
spellingShingle Dipjyoti Das
Dibyendu Das
Ranjith Padinhateeri
Collective force generated by multiple biofilaments can exceed the sum of forces due to individual ones
New Journal of Physics
non-equilibrium chemical switching
dynamics of biofilaments
collective force generation
actin and microtubules
87.16.A-
87.16.Ka
title Collective force generated by multiple biofilaments can exceed the sum of forces due to individual ones
title_full Collective force generated by multiple biofilaments can exceed the sum of forces due to individual ones
title_fullStr Collective force generated by multiple biofilaments can exceed the sum of forces due to individual ones
title_full_unstemmed Collective force generated by multiple biofilaments can exceed the sum of forces due to individual ones
title_short Collective force generated by multiple biofilaments can exceed the sum of forces due to individual ones
title_sort collective force generated by multiple biofilaments can exceed the sum of forces due to individual ones
topic non-equilibrium chemical switching
dynamics of biofilaments
collective force generation
actin and microtubules
87.16.A-
87.16.Ka
url https://doi.org/10.1088/1367-2630/16/6/063032
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AT dibyendudas collectiveforcegeneratedbymultiplebiofilamentscanexceedthesumofforcesduetoindividualones
AT ranjithpadinhateeri collectiveforcegeneratedbymultiplebiofilamentscanexceedthesumofforcesduetoindividualones