Clusiomitidae, A New Family of Eocene Fossil Acalyptratae, with Revision of <i>Acartophthalmites</i> Hennig and <i>Clusiomites</i> Gen. Nov. (Diptera)

The Eocene Baltic amber fossil flies of the genus <i>Acartophthalmites</i> Hennig, 1965 (Diptera: Acalyptratae) are revised. Seven species are recognized and described or redescribed. Five species, <i>A. crassipes</i> sp. nov., <i>A. luridus</i> sp. nov., <i>...

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Bibliographic Details
Main Authors: Jindřich Roháček, Christel Hoffeins
Format: Article
Language:English
Published: MDPI AG 2021-12-01
Series:Insects
Subjects:
Online Access:https://www.mdpi.com/2075-4450/12/12/1123
Description
Summary:The Eocene Baltic amber fossil flies of the genus <i>Acartophthalmites</i> Hennig, 1965 (Diptera: Acalyptratae) are revised. Seven species are recognized and described or redescribed. Five species, <i>A. crassipes</i> sp. nov., <i>A. luridus</i> sp. nov., <i>A. rugosus</i> sp. nov., <i>A. tertiaria</i> Hennig, 1965 (type species) and <i>A. willii</i> Pérez-de la Fuente, Hoffeins et Roháček, 2018 are retained in <i>Acartophthalmites</i> while <i>Clusiomites</i> gen. nov. is described for two other species, <i>C. clusioides</i> (Roháček, 2016) comb. nov. (type species) and <i>C. ornatus</i> sp. nov. Relationships of these fossil taxa are discussed and, because they cannot be confidently placed in any known family of Diptera, a new family, Clusiomitidae, is established for them. Clusiomitidae is recognized as a family of Opomyzoidea, probably most closely allied to Clusiidae. These results again confirmed that the diversity of acalyptrate flies was very high in the Mid-late Eocene amber forest.
ISSN:2075-4450