Systematics of the Rubidgeinae (Therapsida: Gorgonopsia)

The subfamily Rubidgeinae, containing the largest known African gorgonopsians, is thoroughly revised. Rubidgeinae is diagnosed by the absence of a blade-like parasphenoid rostrum and reduction or absence of the preparietal. Seven rubidgeine species from the Karoo Basin of South Africa are recognized...

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Main Author: Christian F. Kammerer
Format: Article
Language:English
Published: PeerJ Inc. 2016-01-01
Series:PeerJ
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Online Access:https://peerj.com/articles/1608.pdf
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author Christian F. Kammerer
author_facet Christian F. Kammerer
author_sort Christian F. Kammerer
collection DOAJ
description The subfamily Rubidgeinae, containing the largest known African gorgonopsians, is thoroughly revised. Rubidgeinae is diagnosed by the absence of a blade-like parasphenoid rostrum and reduction or absence of the preparietal. Seven rubidgeine species from the Karoo Basin of South Africa are recognized as valid: Aelurognathus tigriceps, Clelandina rubidgei, Dinogorgon rubidgei, Leontosaurus vanderhorsti, Rubidgea atrox, Smilesaurus ferox, and Sycosaurus laticeps. Rubidgeines are also present in other African basins: A. tigriceps and S. laticeps occur in the Upper Madumabisa Mudstone Formation of Zambia, and D. rubidgei, R. atrox, and the endemic species Ruhuhucerberus haughtoni comb. nov. and Sycosaurus nowaki comb. nov. occur in the Usili Formation of Tanzania. Aelurognathus nyasaensis from the Chiweta Beds of Malawi also represents a rubidgeine, but of uncertain generic referral pending further preparation. No rubidgeine material is known outside of Africa: the purported Russian rubidgeine Leogorgon klimovensis is not clearly referable to this group and may not be diagnosable. Phylogenetic analysis of rubidgeines reveals strong support for a clade (Rubidgeini) of advanced rubidgeines including Clelandina, Dinogorgon, Leontosaurus, and Rubidgea. Support for Smilesaurus as a rubidgeine is weak; it may, as previous authors have suggested, represent an independent evolution of large body size from an Arctops-like ancestor. Temporally, rubidgeines are restricted to the Late Permian, first appearing in the Tropidostoma Assemblage Zone and reaching highest diversity in the Cistecephalus and Daptocephalus assemblage zones of the Beaufort Group.
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spelling doaj.art-a963d84031904532846596868938b7922023-12-03T11:05:35ZengPeerJ Inc.PeerJ2167-83592016-01-014e160810.7717/peerj.1608Systematics of the Rubidgeinae (Therapsida: Gorgonopsia)Christian F. KammererThe subfamily Rubidgeinae, containing the largest known African gorgonopsians, is thoroughly revised. Rubidgeinae is diagnosed by the absence of a blade-like parasphenoid rostrum and reduction or absence of the preparietal. Seven rubidgeine species from the Karoo Basin of South Africa are recognized as valid: Aelurognathus tigriceps, Clelandina rubidgei, Dinogorgon rubidgei, Leontosaurus vanderhorsti, Rubidgea atrox, Smilesaurus ferox, and Sycosaurus laticeps. Rubidgeines are also present in other African basins: A. tigriceps and S. laticeps occur in the Upper Madumabisa Mudstone Formation of Zambia, and D. rubidgei, R. atrox, and the endemic species Ruhuhucerberus haughtoni comb. nov. and Sycosaurus nowaki comb. nov. occur in the Usili Formation of Tanzania. Aelurognathus nyasaensis from the Chiweta Beds of Malawi also represents a rubidgeine, but of uncertain generic referral pending further preparation. No rubidgeine material is known outside of Africa: the purported Russian rubidgeine Leogorgon klimovensis is not clearly referable to this group and may not be diagnosable. Phylogenetic analysis of rubidgeines reveals strong support for a clade (Rubidgeini) of advanced rubidgeines including Clelandina, Dinogorgon, Leontosaurus, and Rubidgea. Support for Smilesaurus as a rubidgeine is weak; it may, as previous authors have suggested, represent an independent evolution of large body size from an Arctops-like ancestor. Temporally, rubidgeines are restricted to the Late Permian, first appearing in the Tropidostoma Assemblage Zone and reaching highest diversity in the Cistecephalus and Daptocephalus assemblage zones of the Beaufort Group.https://peerj.com/articles/1608.pdfSynapsidaTherapsidaGorgonopsiaPermianAfricaPhylogeny
spellingShingle Christian F. Kammerer
Systematics of the Rubidgeinae (Therapsida: Gorgonopsia)
PeerJ
Synapsida
Therapsida
Gorgonopsia
Permian
Africa
Phylogeny
title Systematics of the Rubidgeinae (Therapsida: Gorgonopsia)
title_full Systematics of the Rubidgeinae (Therapsida: Gorgonopsia)
title_fullStr Systematics of the Rubidgeinae (Therapsida: Gorgonopsia)
title_full_unstemmed Systematics of the Rubidgeinae (Therapsida: Gorgonopsia)
title_short Systematics of the Rubidgeinae (Therapsida: Gorgonopsia)
title_sort systematics of the rubidgeinae therapsida gorgonopsia
topic Synapsida
Therapsida
Gorgonopsia
Permian
Africa
Phylogeny
url https://peerj.com/articles/1608.pdf
work_keys_str_mv AT christianfkammerer systematicsoftherubidgeinaetherapsidagorgonopsia