Mechanism of squalene biosynthesis: evidence against the involvement of free nerolidyl pyrophosphate

Several mechanisms that utilize farnesyl pyrophosphate and nerolidyl pyrophosphate as condensing substrates have been postulated for the asymmetric condensation reaction in squalene biosynthesis. Although there is ample evidence that farnesyl pyrophosphate is a substrate for this reaction, there has...

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Main Authors: S.S. Sofer, H.C. Rilling
Format: Article
Language:English
Published: Elsevier 1969-03-01
Series:Journal of Lipid Research
Subjects:
Online Access:http://www.sciencedirect.com/science/article/pii/S0022227520426665
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author S.S. Sofer
H.C. Rilling
author_facet S.S. Sofer
H.C. Rilling
author_sort S.S. Sofer
collection DOAJ
description Several mechanisms that utilize farnesyl pyrophosphate and nerolidyl pyrophosphate as condensing substrates have been postulated for the asymmetric condensation reaction in squalene biosynthesis. Although there is ample evidence that farnesyl pyrophosphate is a substrate for this reaction, there has been no information concerning the role of nerolidyl pyrophosphate. We have made the following observations that demonstrate that nerolidyl pyrophosphate cannot be a free intermediate in squalene biosynthesis. (a) There is no significant interconversion of farnesyl pyrophosphate and nerolidyl pyrophosphate in a squalene-synthesizing system from yeast. (b) Nerolidyl-1-3H2 phosphate is not converted to squalene in the presence or absence of farnesyl pyrophosphate. (c) The addition of unlabeled nerolidyl pyrophosphate to incubation mixtures does not alter the relative loss of α-hydrogens from farnesyl pyrophosphate during its conversion to squalene.The synthesis of nerolidyl-1-3H2 rophosphate is described. Chromatographic methods for the separation of pyrophosphate esters of triprenols and terpenols are included.
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spelling doaj.art-ab5481d67b95409cbd4aea990a0042872022-12-21T21:57:57ZengElsevierJournal of Lipid Research0022-22751969-03-01102183187Mechanism of squalene biosynthesis: evidence against the involvement of free nerolidyl pyrophosphateS.S. Sofer0H.C. Rilling1Department of Biochemistry, University of Utah College of Medicine, Salt Lake City, Utah 84112Department of Biochemistry, University of Utah College of Medicine, Salt Lake City, Utah 84112Several mechanisms that utilize farnesyl pyrophosphate and nerolidyl pyrophosphate as condensing substrates have been postulated for the asymmetric condensation reaction in squalene biosynthesis. Although there is ample evidence that farnesyl pyrophosphate is a substrate for this reaction, there has been no information concerning the role of nerolidyl pyrophosphate. We have made the following observations that demonstrate that nerolidyl pyrophosphate cannot be a free intermediate in squalene biosynthesis. (a) There is no significant interconversion of farnesyl pyrophosphate and nerolidyl pyrophosphate in a squalene-synthesizing system from yeast. (b) Nerolidyl-1-3H2 phosphate is not converted to squalene in the presence or absence of farnesyl pyrophosphate. (c) The addition of unlabeled nerolidyl pyrophosphate to incubation mixtures does not alter the relative loss of α-hydrogens from farnesyl pyrophosphate during its conversion to squalene.The synthesis of nerolidyl-1-3H2 rophosphate is described. Chromatographic methods for the separation of pyrophosphate esters of triprenols and terpenols are included.http://www.sciencedirect.com/science/article/pii/S0022227520426665yeastfarnesyl pyrophosphatetriprenol and terpenol pyrophosphateschromatographysynthesis of nerolidyl pyrophosphate
spellingShingle S.S. Sofer
H.C. Rilling
Mechanism of squalene biosynthesis: evidence against the involvement of free nerolidyl pyrophosphate
Journal of Lipid Research
yeast
farnesyl pyrophosphate
triprenol and terpenol pyrophosphates
chromatography
synthesis of nerolidyl pyrophosphate
title Mechanism of squalene biosynthesis: evidence against the involvement of free nerolidyl pyrophosphate
title_full Mechanism of squalene biosynthesis: evidence against the involvement of free nerolidyl pyrophosphate
title_fullStr Mechanism of squalene biosynthesis: evidence against the involvement of free nerolidyl pyrophosphate
title_full_unstemmed Mechanism of squalene biosynthesis: evidence against the involvement of free nerolidyl pyrophosphate
title_short Mechanism of squalene biosynthesis: evidence against the involvement of free nerolidyl pyrophosphate
title_sort mechanism of squalene biosynthesis evidence against the involvement of free nerolidyl pyrophosphate
topic yeast
farnesyl pyrophosphate
triprenol and terpenol pyrophosphates
chromatography
synthesis of nerolidyl pyrophosphate
url http://www.sciencedirect.com/science/article/pii/S0022227520426665
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