The Functional Role of Spontaneously Opening GABAA Receptors in Neural Transmission

Ionotropic type of γ-aminobutyric acid receptors (GABAARs) produce two forms of inhibitory signaling: phasic inhibition generated by rapid efflux of neurotransmitter GABA into the synaptic cleft with subsequent binding to GABAARs, and tonic inhibition generated by persistent activation of extrasynap...

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Main Authors: Nathanael O’Neill, Sergiy Sylantyev
Format: Article
Language:English
Published: Frontiers Media S.A. 2019-03-01
Series:Frontiers in Molecular Neuroscience
Subjects:
Online Access:https://www.frontiersin.org/article/10.3389/fnmol.2019.00072/full
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author Nathanael O’Neill
Sergiy Sylantyev
author_facet Nathanael O’Neill
Sergiy Sylantyev
author_sort Nathanael O’Neill
collection DOAJ
description Ionotropic type of γ-aminobutyric acid receptors (GABAARs) produce two forms of inhibitory signaling: phasic inhibition generated by rapid efflux of neurotransmitter GABA into the synaptic cleft with subsequent binding to GABAARs, and tonic inhibition generated by persistent activation of extrasynaptic and/or perisynaptic GABAARs by GABA continuously present in the extracellular space. It is widely accepted that phasic and tonic GABAergic inhibition is mediated by receptor groups of distinct subunit composition and modulated by different cytoplasmic mechanisms. Recently, however, it has been demonstrated that spontaneously opening GABAARs (s-GABAARs), which do not need GABA binding to enter an active state, make a significant input into tonic inhibitory signaling. Due to GABA-independent action mode, s-GABAARs promise new safer options for therapy of neural disorders (such as epilepsy) devoid of side effects connected to abnormal fluctuations of GABA concentration in the brain. However, despite the potentially important role of s-GABAARs in neural signaling, they still remain out of focus of neuroscience studies, to a large extent due to technical difficulties in their experimental research. Here, we summarize present data on s-GABAARs functional properties and experimental approaches that allow isolation of s-GABAARs effects from those of conventional (GABA-dependent) GABAARs.
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spelling doaj.art-d94a1d5e740d44e5922beced84000e402022-12-21T19:52:55ZengFrontiers Media S.A.Frontiers in Molecular Neuroscience1662-50992019-03-011210.3389/fnmol.2019.00072449596The Functional Role of Spontaneously Opening GABAA Receptors in Neural TransmissionNathanael O’NeillSergiy SylantyevIonotropic type of γ-aminobutyric acid receptors (GABAARs) produce two forms of inhibitory signaling: phasic inhibition generated by rapid efflux of neurotransmitter GABA into the synaptic cleft with subsequent binding to GABAARs, and tonic inhibition generated by persistent activation of extrasynaptic and/or perisynaptic GABAARs by GABA continuously present in the extracellular space. It is widely accepted that phasic and tonic GABAergic inhibition is mediated by receptor groups of distinct subunit composition and modulated by different cytoplasmic mechanisms. Recently, however, it has been demonstrated that spontaneously opening GABAARs (s-GABAARs), which do not need GABA binding to enter an active state, make a significant input into tonic inhibitory signaling. Due to GABA-independent action mode, s-GABAARs promise new safer options for therapy of neural disorders (such as epilepsy) devoid of side effects connected to abnormal fluctuations of GABA concentration in the brain. However, despite the potentially important role of s-GABAARs in neural signaling, they still remain out of focus of neuroscience studies, to a large extent due to technical difficulties in their experimental research. Here, we summarize present data on s-GABAARs functional properties and experimental approaches that allow isolation of s-GABAARs effects from those of conventional (GABA-dependent) GABAARs.https://www.frontiersin.org/article/10.3389/fnmol.2019.00072/fullGABA-A receptorGABA-independent inhibitionphasic conductancetonic conductanceG-proteins
spellingShingle Nathanael O’Neill
Sergiy Sylantyev
The Functional Role of Spontaneously Opening GABAA Receptors in Neural Transmission
Frontiers in Molecular Neuroscience
GABA-A receptor
GABA-independent inhibition
phasic conductance
tonic conductance
G-proteins
title The Functional Role of Spontaneously Opening GABAA Receptors in Neural Transmission
title_full The Functional Role of Spontaneously Opening GABAA Receptors in Neural Transmission
title_fullStr The Functional Role of Spontaneously Opening GABAA Receptors in Neural Transmission
title_full_unstemmed The Functional Role of Spontaneously Opening GABAA Receptors in Neural Transmission
title_short The Functional Role of Spontaneously Opening GABAA Receptors in Neural Transmission
title_sort functional role of spontaneously opening gabaa receptors in neural transmission
topic GABA-A receptor
GABA-independent inhibition
phasic conductance
tonic conductance
G-proteins
url https://www.frontiersin.org/article/10.3389/fnmol.2019.00072/full
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