| Summary: | (1) Plants are robust and densely branched; (2) branches are densely foliated and slightly flattened; (3) branch leaves oblong to oblong-obovate, erect-patent, deeply concave, and apices obtuse to truncate, rugose or recurved; (3) leaf margins are incurved, denticulate with simple teeth; (4) lamina cells are slightly prorulate or almost smooth, moderately thin-walled, and sub-apical cells sub-rhomboidal and thick-walled; (5) inner perichaetial leaves are oblong, strongly laciniate with obtuse or even truncate apices; (6) setae are 8 - 9 mm long, papillose above, papillae low, smooth half below.
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