Taxonomy, geographic variation and population genetics of Bornean and Sumatran orangutans by Goossens, Benoît, Chikhi, Lounès, Mohd. Fairus Jalil, James, Sheena, Lackman-Ancrenaz, Isabelle, Bruford, Michael W.

“…Moreover, three subspecies have been described on Borneo Island: P. p. pygmaeus in Sarawak and west Kalimantan, P. p. morio in Sabah and east Kalimantan and P. p. wurmbii in central and south Kalimantan. Despite this, little is known about the intra-subspecific variation between isolated Bornean populations and among the Sumatran populations. …”
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