Land-use changes, farm management and the decline of butterflies associated with semi-natural grasslands in southern Sweden by Sven G. Nilsson, Markus Franzén, Lars Pettersson

“…Many typical grassland species that were numerous in former times have declined severely; among those Hesperia comma, Lycaena virgaureae, Lycaena hippothoe, Argynnis adippe, and Polyommatus semiargus. …”
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