Subspecies of Hypolepis rugosula (Dennstaedtiaceae; Pteridophyta) around the world: morphological and biogeographic perspectives by Pedro Bond Schwartsburd, Jefferson Prado

“…The "Hypolepis rugosula complex" has been the subject of great debate among pteridologists: some have considered H. rugosula a single subcosmopolitan (or circum-Antarctic) species, whereas others have considered it a species-complex, encompassing several species. …”
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