IgM Antibodies Can Access Cryptic Antigens Denied to IgG: Hypothesis on Novel Binding Mechanism

Antibodies are well-known protein mediators of immunity. IgM is the primordial member and the neglected sibling of the later-evolved and more proficient IgG in regard to their therapeutic and diagnostic use. Serendipitously, however, we found a paradox: While murine IgM antibodies specific for guano...

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Main Authors: Eric Chun Yiu Law, Danny Tze Ming Leung, Frankie Chi Hang Tam, Kitty Kit Ting Cheung, Naomi Hua Yin Cheng, Pak Leong Lim
Format: Article
Language:English
Published: Frontiers Media S.A. 2019-08-01
Series:Frontiers in Immunology
Subjects:
Online Access:https://www.frontiersin.org/article/10.3389/fimmu.2019.01820/full
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author Eric Chun Yiu Law
Danny Tze Ming Leung
Frankie Chi Hang Tam
Frankie Chi Hang Tam
Kitty Kit Ting Cheung
Naomi Hua Yin Cheng
Pak Leong Lim
Pak Leong Lim
author_facet Eric Chun Yiu Law
Danny Tze Ming Leung
Frankie Chi Hang Tam
Frankie Chi Hang Tam
Kitty Kit Ting Cheung
Naomi Hua Yin Cheng
Pak Leong Lim
Pak Leong Lim
author_sort Eric Chun Yiu Law
collection DOAJ
description Antibodies are well-known protein mediators of immunity. IgM is the primordial member and the neglected sibling of the later-evolved and more proficient IgG in regard to their therapeutic and diagnostic use. Serendipitously, however, we found a paradox: While murine IgM antibodies specific for guanosine triphosphate (GTP) were able to recognize native guanylyl antigens found in primate or rat muscle tissues by immunofluorescence assays (which mimicked the auto-antibodies from autoimmune patients to skeletal or smooth muscle), the murine and human IgG counterparts failed. The results were replicated in cell-free direct binding assays using small latex microspheres decorated densely with GTP. The IgG antibodies could bind, however, if GTP was presented more spaciously on larger particles or as a univalent hapten. Accordingly, oligomerization of GTP (30-mer) destroyed the binding of the IgG antibodies but enhanced that of the IgMs in inhibition ELISA. We reason that, contrary to current belief, IgM does not bind in a lock-and-key manner like IgG. We hypothesize that whereas the intact and rigid antigen-binding site of IgG hinders the antibody from docking with antigens that are obstructed, in IgM, the two component polypeptides of the antigen-binding site can dissociate from each other and navigate individually through obstacles like the ancestral single-polypeptide antibodies found in sharks and camelids, both components eventually re-grouping around the antigen. We further speculate that polyreactive IgMs, which enigmatically bind to more than one type of antigen, use the same modus operandi. These findings call for a re-look at the clinical potential of IgM antibodies particularly in specific areas of cancer therapy, tissue pathology and vaccine design, where IgG antibodies have failed due to target inaccessibility.
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spelling doaj.art-12e4da23223a4cdfb4efefda72c385482022-12-22T00:02:42ZengFrontiers Media S.A.Frontiers in Immunology1664-32242019-08-011010.3389/fimmu.2019.01820468889IgM Antibodies Can Access Cryptic Antigens Denied to IgG: Hypothesis on Novel Binding MechanismEric Chun Yiu Law0Danny Tze Ming Leung1Frankie Chi Hang Tam2Frankie Chi Hang Tam3Kitty Kit Ting Cheung4Naomi Hua Yin Cheng5Pak Leong Lim6Pak Leong Lim7Clinical Immunology Unit, The Chinese University of Hong Kong, Hong Kong, ChinaClinical Immunology Unit, The Chinese University of Hong Kong, Hong Kong, ChinaClinical Immunology Unit, The Chinese University of Hong Kong, Hong Kong, ChinaIgGENE, FoTan, Hong Kong, ChinaClinical Immunology Unit, The Chinese University of Hong Kong, Hong Kong, ChinaClinical Immunology Unit, The Chinese University of Hong Kong, Hong Kong, ChinaClinical Immunology Unit, The Chinese University of Hong Kong, Hong Kong, ChinaIgGENE, FoTan, Hong Kong, ChinaAntibodies are well-known protein mediators of immunity. IgM is the primordial member and the neglected sibling of the later-evolved and more proficient IgG in regard to their therapeutic and diagnostic use. Serendipitously, however, we found a paradox: While murine IgM antibodies specific for guanosine triphosphate (GTP) were able to recognize native guanylyl antigens found in primate or rat muscle tissues by immunofluorescence assays (which mimicked the auto-antibodies from autoimmune patients to skeletal or smooth muscle), the murine and human IgG counterparts failed. The results were replicated in cell-free direct binding assays using small latex microspheres decorated densely with GTP. The IgG antibodies could bind, however, if GTP was presented more spaciously on larger particles or as a univalent hapten. Accordingly, oligomerization of GTP (30-mer) destroyed the binding of the IgG antibodies but enhanced that of the IgMs in inhibition ELISA. We reason that, contrary to current belief, IgM does not bind in a lock-and-key manner like IgG. We hypothesize that whereas the intact and rigid antigen-binding site of IgG hinders the antibody from docking with antigens that are obstructed, in IgM, the two component polypeptides of the antigen-binding site can dissociate from each other and navigate individually through obstacles like the ancestral single-polypeptide antibodies found in sharks and camelids, both components eventually re-grouping around the antigen. We further speculate that polyreactive IgMs, which enigmatically bind to more than one type of antigen, use the same modus operandi. These findings call for a re-look at the clinical potential of IgM antibodies particularly in specific areas of cancer therapy, tissue pathology and vaccine design, where IgG antibodies have failed due to target inaccessibility.https://www.frontiersin.org/article/10.3389/fimmu.2019.01820/fullIgMantibody specificityguanosine triphosphatelock-and-keyantigen-binding sitesteric hindrance
spellingShingle Eric Chun Yiu Law
Danny Tze Ming Leung
Frankie Chi Hang Tam
Frankie Chi Hang Tam
Kitty Kit Ting Cheung
Naomi Hua Yin Cheng
Pak Leong Lim
Pak Leong Lim
IgM Antibodies Can Access Cryptic Antigens Denied to IgG: Hypothesis on Novel Binding Mechanism
Frontiers in Immunology
IgM
antibody specificity
guanosine triphosphate
lock-and-key
antigen-binding site
steric hindrance
title IgM Antibodies Can Access Cryptic Antigens Denied to IgG: Hypothesis on Novel Binding Mechanism
title_full IgM Antibodies Can Access Cryptic Antigens Denied to IgG: Hypothesis on Novel Binding Mechanism
title_fullStr IgM Antibodies Can Access Cryptic Antigens Denied to IgG: Hypothesis on Novel Binding Mechanism
title_full_unstemmed IgM Antibodies Can Access Cryptic Antigens Denied to IgG: Hypothesis on Novel Binding Mechanism
title_short IgM Antibodies Can Access Cryptic Antigens Denied to IgG: Hypothesis on Novel Binding Mechanism
title_sort igm antibodies can access cryptic antigens denied to igg hypothesis on novel binding mechanism
topic IgM
antibody specificity
guanosine triphosphate
lock-and-key
antigen-binding site
steric hindrance
url https://www.frontiersin.org/article/10.3389/fimmu.2019.01820/full
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